| Strain | ||||
|---|---|---|---|---|
| Urease | Negative | Negative | Negative | Positive |
| Mannitol assimilation | Negative | Positive | Positive | Positive |
| Maltose assimilation | Negative | Positive | Positive | Positive |
| Adipate assimilation | Negative | Positive | Negative | Negative |
| Citrate assimilation | Positive | Positive | Negative | Positive |